Genetics and Evolution

Molecular Basis of Inheritance - Function of 5’ Capping


Introduction to Molecular Basis of Inheritance


Structure and Function of DNA and RNA


Gene Expression and Regulation


Central Dogma of Molecular Biology


Transcription


Translation


Regulation of Gene Expression


Function of 5’ Capping


Mechanism of 5’ Capping

  1. RNA polymerase synthesizes pre-mRNA.
  2. Addition of a cap structure to the 5’ end.
  3. Cap structure includes 7-methylguanosine, methylation, and linkage.
  4. Cap binding proteins and enzymes recognize the cap structure.

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Significance of 5’ Capping


Importance of Exonuclease Protection


Stability Enhancement of mRNA


Facilitation of Nuclear Export


Promotion of Translation Initiation


Regulation of Gene Expression


Aberrations in 5’ Capping


Examples of 5’ Capping in Gene Expression


5’ Capping and Transcription Start Site


Conclusion

Examples of 5’ Capping


5’ Capping and Translation Efficiency


5’ Capping and Protein Synthesis Rates


5’ Capping Defects and Disease


Experimental Techniques to Study 5’ Capping


Future Perspectives


Summary


References

  1. Shatkin, A. J. (1976). Capping of eucaryotic mRNAs. Cell, 9(4 Pt 2), 645–653. doi: 10.1016/0092-8674(76)90024-0

  2. Furuichi, Y., & Shatkin, A. J. (2000). Viral and cellular mRNA capping: Past and prospects. Advances in virus research, 55, 135–184. doi: 10.1016/S0065-3527(00)55004-X

  3. Vazquez-Pianzola, P., Adamson, M. E., & Capraro, F. D. (2020). Distinct ribosome populations preferentially translate either early or late-occurring mRNAs in Drosophila melanogaster embryos. eLife, 9, e63184. doi: 10.7554/eLife.63184


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