Ecosystem
Decomposition
You may have heard of the earthworm being referred to as the farmer’s ‘friend’
This is so because they help in the breakdown of complex organic matter as well as in loosening of the soil
Similarly, decomposers break down complex organic matter into inorganic substances like carbon dioxide, water and nutrients and the process is called decomposition
Dead plant remains such as leaves, bark, flowers and dead remains of animals, including fecal matter, constitute detritus, which is the raw material for decomposition
The important steps in the process of decomposition are fragmentation, leaching, catabolism, humification and mineralisation
Detritivores (e.g., earthworm) break down detritus into smaller particles
This process is called fragmentation
By the process of leaching, watersoluble inorganic nutrients go down into the soil horizon and get precipitated as unavailable salts
Bacterial and fungal enzymes degrade detritus into simpler inorganic substances
This process is called as catabolism
It is important to note that all the above steps in decomposition operate simultaneously on the detritus (Figure 12.1)
Humification and mineralisation occur during decomposition in the soil
Humification leads to accumulation of a dark coloured amorphous substance called humus that is highly resistant to microbial action and undergoes decomposition at an extremely slow rate
Being colloidal in nature it serves as a reservoir of nutrients
The humus is further degraded by some microbes and release of inorganic nutrients occur by the process known as mineralisation
Decomposition is largely an oxygen-requiring process
The rate of decomposition is controlled by chemical composition of detritus and climatic factors
In a particular climatic condition, decomposition rate is slower if detritus is rich in lignin and chitin, and quicker, if detritus is rich in nitrogen and water-soluble substances like sugars
Temperature and soil moisture are the most important climatic factors that regulate decomposition through their effects on the activities of soil microbes
Energy Flow
Except for the deep sea hydro-thermal ecosystem, sun is the only source of energy for all ecosystems on Earth
Of the incident solar radiation less than 50 per cent of it is photosynthetically active radiation (PAR)
We know that plants and photosynthetic bacteria (autotrophs), fix Sun’s radiant energy to make food from simple inorganic materials
Plants capture only 2-10 per cent of the PAR and this small amount of energy sustains the entire living world
So, it is very important to know how the solar energy captured by plants flows through different organisms of an ecosystem
All organisms are dependent for their food on producers, either directly or indirectly
So you find unidirectional flow of energy from the sun to producers and then to consumers
Is this in keeping with the first law of thermodynamics?
Further, ecosystems are not exempt from the Second Law of thermodynamics
They need a constant supply of energy to synthesise the molecules they require, to counteract the universal tendency toward increasing disorderliness
The green plant in the ecosystem are called producers
In a terrestrial ecosystem, major producers are herbaceous and woody plants
Likewise, producers in an aquatic ecosystem are various species like phytoplankton, algae and higher plants
You have read about the food chains and webs that exist in nature
Starting from the plants (or producers) food chains or rather webs are formed such that an animal feeds on a plant or on another animal and in turn is food for another
The chain or web is formed because of this interdependency
No energy that is trapped into an organism remains in it for ever
The energy trapped by the producer, hence, is either passed on to a consumer or the organism dies
Death of organism is the beginning of the detritus food chain/web
All animals depend on plants (directly or indirectly) for their food needs
They are hence called consumers and also heterotrophs
If they feed on the producers, the plants, they are called primary consumers, and if the animals eat other animals which in turn eat the plants (or their produce) they are called secondary consumers
Likewise, you could have tertiary consumers too
Obviously the primary consumers will be herbivores
Some common herbivores are insects, birds and mammals in terrestrial ecosystem and molluscs in aquatic ecosystem
The consumers that feed on these herbivores are carnivores, or more correctly primary carnivores (though secondary consumers)
Those animals that depend on the primary carnivores for food are labelled secondary carnivores
$\underset{(Producer)}{Grass}\rightarrow$ $\underset{\text{(Primary consumer)}}{Goat}\rightarrow$ $\underset{\text{(Secondary consumer)}}{Man}\rightarrow$
The detritus food chain (DFC) begins with dead organic matter
It is made up of decomposers which are heterotrophic organisms, mainly fungi and bacteria
They meet their energy and nutrient requirements by degrading dead organic matter or detritus
These are also known as saprotrophs (sapro: to decompose)
Decomposers secrete digestive enzymes that breakdown dead and waste materials into simple, inorganic materials, which are subsequently absorbed by them
In an aquatic ecosystem, GFC is the major conduit for energy flow
As against this, in a terrestrial ecosystem, a much larger fraction of energy flows through the detritus food chain than through the GFC
Detritus food chain may be connected with the grazing food chain at some levels: some of the organisms of DFC are prey to the GFC animals, and in a natural ecosystem, some animals like cockroaches, crows, etc., are omnivores
These natural interconnection of food chains make it a food web
How would you classify human beings! Organisms occupy a place in the natural surroundings or in a community according to their feeding relationship with other organisms
Based on the source of their nutrition or food, organisms occupy a specific place in the food chain that is known as their trophic level
The important point to note is that the amount of energy decreases at successive trophic levels
When any organism dies it is converted to detritus or dead biomass that serves as an energy source for decomposers
Organisms at each trophic level depend on those at the lower trophic level for their energy demands
Each trophic level has a certain mass of living material at a particular time called as the standing crop
The standing crop is measured as the mass of living organisms (biomass) or the number in a unit area
The biomass of a species is expressed in terms of fresh or dry weight
Measurement of biomass in terms of dry weight is more accurate. Why?
The number of trophic levels in the grazing food chain is restricted as the transfer of energy follows 10 per cent law – only 10 per cent of the energy is transferred to each trophic level from the lower trophic level
In nature, it is possible to have so many levels – producer, herbivore, primary carnivore, secondary carnivore in the grazing food chain (Figure 12.3)
Do you think there is any such limitation in a detritus food chain?